:root {
  color-scheme: light;
  --ink: #14261C;
  --forest: #1A3328;
  --sage: #D8E2D4;
  --cream: #F5F0E6;
  --copper: #B56A3A;
  --coral: #C46B55;
  --white: #FFFEFB;
  --muted: rgba(20, 38, 28, 0.72);
  --line: rgba(20, 38, 28, 0.14);
  --paper: var(--cream);
  --soft: var(--sage);
  --moss: var(--forest);
  --moss-dark: var(--forest);
  --clay: var(--copper);
  --gold: var(--copper);
  --water: var(--forest);
  --shadow: 0 16px 40px rgba(20, 38, 28, 0.1);
  --font-sans: Inter, system-ui, -apple-system, BlinkMacSystemFont, "Segoe UI", sans-serif;
  --font-serif: "Source Serif 4", "Iowan Old Style", "Palatino Linotype", Palatino, Georgia, serif;
}

* {
  box-sizing: border-box;
}

html {
  scroll-behavior: smooth;
}

body {
  margin: 0;
  font-family: var(--font-sans);
  background: var(--paper);
  color: var(--ink);
}

h1,
h2,
h3,
.brand-text {
  font-family: var(--font-serif);
  font-weight: 600;
  letter-spacing: -0.02em;
}

a {
  color: inherit;
}

.site-header {
  position: sticky;
  top: 0;
  z-index: 20;
  display: flex;
  align-items: center;
  justify-content: space-between;
  gap: 24px;
  min-height: 88px;
  padding: 14px clamp(18px, 4vw, 56px);
  border-bottom: 1px solid var(--line);
  background: rgba(245, 240, 230, 0.92);
  backdrop-filter: blur(16px);
}

.brand {
  display: inline-flex;
  align-items: center;
  gap: 12px;
  text-decoration: none;
  white-space: nowrap;
}

.brand-mark {
  display: block;
  width: 48px;
  height: 48px;
  flex: 0 0 auto;
  object-fit: contain;
}

.brand-text {
  font-size: clamp(1.35rem, 2.6vw, 1.85rem);
  font-weight: 600;
  line-height: 1;
  color: var(--ink);
}

.site-nav {
  display: flex;
  align-items: center;
  gap: clamp(18px, 3vw, 34px);
}

.site-nav a {
  position: relative;
  padding: 10px 0;
  color: var(--muted);
  font-size: 0.92rem;
  font-weight: 700;
  letter-spacing: 0.01em;
  text-decoration: none;
}

.site-nav a:hover,
.site-nav a:focus-visible,
.site-nav a[aria-current="page"] {
  color: var(--ink);
}

.site-nav a[aria-current="page"]::after {
  position: absolute;
  right: 0;
  bottom: 3px;
  left: 0;
  height: 2px;
  background: var(--copper);
  content: "";
}

.eyebrow {
  margin: 0 0 10px;
  color: var(--gold);
  font-family: var(--font-sans);
  font-size: 0.78rem;
  font-weight: 800;
  letter-spacing: 0.06em;
  text-transform: uppercase;
}

.button {
  display: inline-flex;
  align-items: center;
  justify-content: center;
  min-height: 48px;
  padding: 0 20px;
  border: 1px solid transparent;
  border-radius: 999px;
  font-family: var(--font-sans);
  font-weight: 700;
  text-decoration: none;
  transition: transform 160ms ease, box-shadow 160ms ease, background-color 160ms ease;
}

.button:hover,
.button:focus-visible {
  transform: translateY(-1px);
}

.button.primary {
  background: var(--copper);
  border-color: var(--copper);
  color: var(--white);
  box-shadow: 0 10px 24px rgba(181, 106, 58, 0.28);
}

.button.primary:hover,
.button.primary:focus-visible {
  background: #a45e32;
  border-color: #a45e32;
}

.section {
  padding: clamp(28px, 5vw, 64px) clamp(18px, 4vw, 56px);
}

.home-articles {
  display: flex;
  align-items: end;
  justify-content: space-between;
  gap: clamp(28px, 6vw, 80px);
  padding: clamp(52px, 8vw, 96px) clamp(18px, 7vw, 110px);
  background:
    radial-gradient(circle at 90% 10%, rgba(181, 106, 58, 0.38), transparent 36%),
    radial-gradient(circle at 12% 88%, rgba(216, 226, 212, 0.12), transparent 28%),
    linear-gradient(145deg, #14261C 0%, var(--forest) 55%, #243F32 100%);
  color: var(--white);
}

.home-articles > div {
  max-width: 850px;
}

.home-articles h1 {
  max-width: 780px;
  margin: 0;
  font-size: clamp(2.8rem, 6.5vw, 4.35rem);
  letter-spacing: -0.03em;
  line-height: 1.02;
}

.home-articles p {
  max-width: 680px;
  margin: 24px 0 0;
  color: rgba(255, 254, 251, 0.78);
  font-size: clamp(1.05rem, 2vw, 1.28rem);
  line-height: 1.65;
}

.home-articles .button {
  flex: 0 0 auto;
}

.home-articles .button.primary {
  background: var(--cream);
  border-color: var(--cream);
  color: var(--forest);
  box-shadow: 0 12px 28px rgba(0, 0, 0, 0.18);
}

.home-articles .button.primary:hover,
.home-articles .button.primary:focus-visible {
  background: var(--white);
  border-color: var(--white);
}

.article-hero {
  padding: clamp(52px, 7vw, 96px) clamp(18px, 7vw, 110px);
  background:
    radial-gradient(circle at 85% 15%, rgba(181, 106, 58, 0.36), transparent 32%),
    linear-gradient(145deg, #14261C, var(--forest));
  color: var(--white);
}

.article-hero-inner,
.article-body {
  max-width: 900px;
  margin: 0 auto;
}

.article-hero .eyebrow {
  color: #e8c4a0;
}

.article-hero h1 {
  max-width: 850px;
  margin: 0;
  font-size: clamp(3rem, 8vw, 4.5rem);
  line-height: 0.96;
}

.article-deck {
  max-width: 720px;
  margin: 28px 0 0;
  color: rgba(255, 255, 255, 0.78);
  font-size: clamp(1.1rem, 2vw, 1.35rem);
  line-height: 1.6;
}

.article-body {
  padding: clamp(48px, 8vw, 100px) clamp(18px, 4vw, 40px);
}

.article-body > p {
  margin: 0 0 28px;
  color: var(--muted);
  font-size: clamp(1.05rem, 1.8vw, 1.2rem);
  line-height: 1.8;
}

.article-body h2 {
  margin: clamp(48px, 7vw, 80px) 0 24px;
  font-size: clamp(2rem, 4vw, 3.25rem);
}

.takeaway-list {
  display: grid;
  gap: 1px;
  margin: 0;
  padding: 1px;
  background: var(--line);
  list-style: none;
}

.takeaway-list li {
  padding: 22px 24px;
  background: var(--white);
  color: var(--muted);
  line-height: 1.6;
}

.takeaway-list strong {
  color: var(--ink);
}

.safety-callout {
  margin-top: clamp(40px, 7vw, 72px);
  padding: clamp(26px, 5vw, 48px);
  border-left: 6px solid var(--copper);
  background: var(--sage);
}

.safety-callout p {
  margin: 0;
  font-size: clamp(1.05rem, 2vw, 1.25rem);
  line-height: 1.7;
}

.section-heading {
  margin-bottom: 24px;
}

.section-heading h2 {
  margin: 0;
  font-size: clamp(2rem, 4vw, 3.25rem);
}

.source-list {
  display: grid;
  gap: 12px;
  margin: 0;
  padding-left: 20px;
  color: var(--muted);
  line-height: 1.55;
}

.source-list a {
  color: var(--moss-dark);
  font-weight: 800;
}

.species-intro {
  max-width: 820px;
  margin-bottom: clamp(32px, 5vw, 56px);
}

.species-intro h1,
.species-intro h2 {
  margin: 0;
  font-size: clamp(2.75rem, 7vw, 4.5rem);
  line-height: 0.98;
  letter-spacing: -0.045em;
}

.species-intro p {
  max-width: 720px;
  margin: 22px 0 0;
  color: var(--muted);
  font-size: clamp(1.05rem, 2vw, 1.3rem);
  line-height: 1.6;
}

.species-filter {
  display: grid;
  gap: clamp(16px, 2.5vw, 22px);
  max-width: 1120px;
  margin: 0 0 clamp(22px, 3vw, 32px);
  padding: clamp(18px, 3vw, 28px);
  border: 1px solid var(--line);
  border-radius: 16px;
  background: var(--white);
  box-shadow: 0 10px 28px rgba(20, 38, 28, 0.05);
}

.filter-intro h3 {
  margin: 0 0 8px;
  font-size: clamp(1.25rem, 2.4vw, 1.55rem);
}

.filter-intro p {
  max-width: 48rem;
  margin: 0;
  color: var(--muted);
  line-height: 1.6;
}

.filter-group {
  margin: 0;
  padding: 0;
  border: 0;
}

.filter-group legend {
  margin-bottom: 10px;
  font-size: 0.86rem;
  font-weight: 800;
}

.filter-hint {
  color: var(--muted);
  font-weight: 600;
}

.filter-chip-row {
  display: flex;
  flex-wrap: wrap;
  gap: 8px;
}

.filter-chip {
  display: inline-flex;
  align-items: center;
  gap: 0;
  cursor: pointer;
}

.filter-chip input {
  position: absolute;
  opacity: 0;
  pointer-events: none;
}

.filter-chip span {
  display: inline-flex;
  align-items: center;
  min-height: 36px;
  padding: 0 14px;
  border: 1px solid var(--line);
  border-radius: 999px;
  background: var(--paper);
  color: var(--muted);
  font-size: 0.88rem;
  font-weight: 700;
  transition: background-color 140ms ease, border-color 140ms ease, color 140ms ease;
}

.filter-chip:hover span,
.filter-chip:focus-within span {
  border-color: rgba(181, 106, 58, 0.45);
  color: var(--ink);
}

.filter-chip input:checked + span {
  border-color: var(--copper);
  background: rgba(181, 106, 58, 0.12);
  color: var(--ink);
}

.filter-chip input:focus-visible + span {
  outline: 3px solid rgba(181, 106, 58, 0.28);
  outline-offset: 2px;
}

.filter-actions {
  display: flex;
  flex-wrap: wrap;
  gap: 10px;
}

.button.subtle {
  min-height: 40px;
  padding: 0 14px;
  border: 1px solid var(--line);
  border-radius: 999px;
  background: transparent;
  color: var(--ink);
  box-shadow: none;
  font: inherit;
  font-weight: 700;
  cursor: pointer;
}

.button.subtle:hover,
.button.subtle:focus-visible {
  border-color: var(--copper);
  background: rgba(181, 106, 58, 0.08);
  transform: none;
}

.species-match-note {
  max-width: 52rem;
  margin: 0 0 18px;
  padding: 14px 16px;
  border-left: 4px solid var(--copper);
  border-radius: 0 10px 10px 0;
  background: rgba(216, 226, 212, 0.55);
  color: var(--muted);
  line-height: 1.55;
}

.species-search {
  display: grid;
  gap: 10px;
  max-width: 820px;
  margin: 0 0 clamp(28px, 4vw, 44px);
}

.species-search label {
  font-size: 0.9rem;
  font-weight: 800;
}

.species-search-row {
  display: grid;
  grid-template-columns: minmax(0, 1fr) auto;
  align-items: center;
  gap: 14px;
}

.species-search input {
  width: 100%;
  min-height: 54px;
  padding: 0 18px;
  border: 1.5px solid var(--line);
  border-radius: 10px;
  background: var(--white);
  color: var(--ink);
  font: inherit;
  font-size: 1rem;
  transition: border-color 160ms ease, box-shadow 160ms ease;
}

.species-search input::placeholder {
  color: rgba(20, 38, 28, 0.48);
}

.species-search input:focus {
  border-color: var(--copper);
  box-shadow: 0 0 0 4px rgba(181, 106, 58, 0.16);
  outline: none;
}

.species-search output {
  min-width: 82px;
  color: var(--muted);
  font-size: 0.9rem;
  font-weight: 700;
  text-align: right;
}

.species-empty-state {
  margin: 0;
  padding: clamp(32px, 6vw, 72px) 24px;
  border: 1px dashed var(--line);
  border-radius: 8px;
  color: var(--muted);
  font-size: 1.05rem;
  text-align: center;
}

.species-grid {
  display: grid;
  grid-template-columns: repeat(4, minmax(0, 1fr));
  gap: 16px;
}

.snake-card {
  display: grid;
  grid-template-rows: auto 1fr;
  overflow: hidden;
  border: 1px solid var(--line);
  border-radius: 12px;
  background: var(--white);
  box-shadow: 0 8px 24px rgba(20, 38, 28, 0.06);
  transition: transform 180ms ease, box-shadow 180ms ease;
}

.snake-card:hover,
.snake-card:focus-within {
  transform: translateY(-2px);
  box-shadow: 0 14px 32px rgba(20, 38, 28, 0.1);
}

.snake-card[hidden] {
  display: none;
}

.card-image-link {
  display: block;
  padding: 10px;
  background: linear-gradient(180deg, rgba(216, 226, 212, 0.85), var(--soft));
}

.card-image-link picture {
  display: block;
  aspect-ratio: 4 / 3;
}

.profile-figure picture {
  display: block;
}

.snake-photo {
  display: block;
  width: 100%;
  height: 100%;
  object-fit: contain;
  border-radius: 6px;
}

.card-body {
  display: grid;
  gap: 8px;
  padding: 16px 16px 18px;
}

.card-body h3 {
  display: flex;
  flex-wrap: wrap;
  align-items: baseline;
  gap: 8px;
  margin: 0;
  font-size: 1.12rem;
  line-height: 1.25;
}

.card-body h3 a {
  text-decoration: none;
}

.card-scientific {
  margin: 0;
  color: var(--muted);
  font-size: 0.84rem;
  font-style: italic;
  line-height: 1.35;
}

.card-match-reasons {
  margin: 0;
  color: var(--forest);
  font-size: 0.8rem;
  font-weight: 700;
  line-height: 1.4;
}

.card-compare {
  display: inline-flex;
  align-items: center;
  gap: 8px;
  margin-top: 4px;
  color: var(--muted);
  font-size: 0.82rem;
  font-weight: 700;
  cursor: pointer;
}

.card-compare input {
  width: 15px;
  height: 15px;
  accent-color: var(--copper);
}

.species-compare {
  margin: 0 0 clamp(24px, 4vw, 36px);
  padding: clamp(18px, 3vw, 28px);
  border: 1px solid var(--line);
  border-radius: 16px;
  background: var(--white);
  box-shadow: 0 10px 28px rgba(20, 38, 28, 0.05);
}

.species-compare-header {
  display: flex;
  flex-wrap: wrap;
  align-items: center;
  justify-content: space-between;
  gap: 12px;
  margin-bottom: 18px;
}

.species-compare-header h3 {
  margin: 0;
  font-size: 1.25rem;
}

.species-compare-grid {
  display: grid;
  grid-template-columns: repeat(3, minmax(0, 1fr));
  gap: 16px;
}

.compare-card {
  display: grid;
  gap: 10px;
  align-content: start;
  padding: 14px;
  border: 1px solid var(--line);
  border-radius: 12px;
  background: var(--paper);
}

.compare-image-link {
  display: block;
  overflow: hidden;
  border-radius: 8px;
  background: var(--soft);
}

.compare-photo {
  display: block;
  width: 100%;
  aspect-ratio: 4 / 3;
  object-fit: cover;
}

.compare-card h3 {
  display: flex;
  flex-wrap: wrap;
  align-items: center;
  gap: 8px;
  margin: 0;
  font-size: 1.05rem;
  line-height: 1.25;
}

.compare-card h3 a {
  text-decoration: none;
}

.compare-card ul {
  margin: 0;
  padding-left: 1.1rem;
  color: var(--muted);
  font-size: 0.88rem;
  line-height: 1.5;
}

.compare-note {
  margin: 0;
  color: var(--muted);
  font-size: 0.9rem;
  line-height: 1.5;
}

.species-noscript {
  margin-top: 18px;
  color: var(--muted);
  line-height: 1.55;
}

.venomous-label {
  display: inline-flex;
  align-items: center;
  margin: 0;
  padding: 3px 8px;
  border-radius: 999px;
  background: rgba(196, 107, 85, 0.14);
  color: var(--coral);
  font-family: var(--font-sans);
  font-size: 0.68rem;
  font-weight: 800;
  letter-spacing: 0.04em;
  line-height: 1.2;
  text-transform: uppercase;
}

.species-profile {
  background: var(--paper);
}

.profile-hero {
  display: grid;
  grid-template-columns: minmax(0, 0.95fr) minmax(320px, 1.05fr);
  gap: clamp(28px, 5vw, 72px);
  align-items: end;
  padding: clamp(40px, 5vw, 72px) clamp(18px, 4vw, 56px);
  background:
    linear-gradient(105deg, rgba(20, 38, 28, 0.94), rgba(26, 51, 40, 0.55)),
    linear-gradient(135deg, var(--accent, var(--copper)), var(--sage));
  color: var(--white);
}

.profile-copy {
  max-width: 720px;
}

.profile-hero .eyebrow {
  color: #e8c4a0;
  margin-bottom: 8px;
}

.profile-copy h1 {
  margin: 8px 0 0;
  font-size: clamp(2.6rem, 6.5vw, 4.2rem);
  line-height: 1.02;
}

.scientific.large {
  margin-top: 12px;
  color: rgba(255, 254, 251, 0.78);
  font-family: var(--font-serif);
  font-size: clamp(1.05rem, 2vw, 1.35rem);
  font-style: italic;
  font-weight: 500;
}

.profile-summary {
  max-width: 38rem;
  margin: 22px 0 0;
  color: rgba(255, 254, 251, 0.88);
  font-size: clamp(1.02rem, 1.7vw, 1.18rem);
  line-height: 1.65;
}

.profile-safety-note {
  max-width: 36rem;
  margin: 18px 0 0;
  padding: 12px 14px;
  border-left: 3px solid rgba(232, 196, 160, 0.9);
  background: rgba(0, 0, 0, 0.18);
  color: rgba(255, 254, 251, 0.9);
  font-size: 0.92rem;
  line-height: 1.5;
}

.profile-figure {
  margin: 0;
}

.not-found {
  min-height: calc(100vh - 76px);
  padding: clamp(56px, 8vw, 96px) clamp(18px, 4vw, 56px);
  background: var(--moss-dark);
  color: var(--white);
}

.not-found h1 {
  max-width: 760px;
  margin: 12px 0 32px;
  font-size: clamp(3rem, 8vw, 4.5rem);
  line-height: 0.95;
}

.profile-figure img {
  display: block;
  width: 100%;
  height: auto;
  aspect-ratio: var(--image-aspect, auto);
  max-height: 620px;
  object-fit: cover;
  border-radius: 12px;
  box-shadow: var(--shadow);
}

.profile-figure figcaption {
  margin-top: 10px;
  color: rgba(255, 255, 255, 0.74);
  font-size: 0.82rem;
}

.profile-figure a {
  color: var(--white);
}

.profile-section {
  padding: clamp(40px, 6vw, 72px) 0;
  border-top: 1px solid var(--line);
}

.profile-section:nth-child(even) {
  background: rgba(216, 226, 212, 0.28);
}

.profile-section-inner {
  max-width: 1120px;
  margin: 0 auto;
  padding: 0 clamp(18px, 4vw, 56px);
}

.profile-section h2 {
  margin: 0 0 clamp(18px, 3vw, 28px);
  font-size: clamp(1.85rem, 3.5vw, 2.75rem);
  letter-spacing: -0.03em;
  line-height: 1.1;
}

.profile-section h3 {
  margin: 28px 0 12px;
  font-family: var(--font-serif);
  font-size: 1.15rem;
  font-weight: 600;
}

.profile-lead,
.profile-section p {
  max-width: 46rem;
  margin: 0 0 16px;
  color: var(--muted);
  font-size: clamp(1.02rem, 1.5vw, 1.12rem);
  line-height: 1.75;
}

.profile-muted {
  color: var(--muted);
}

.profile-fact-grid {
  display: grid;
  grid-template-columns: repeat(3, minmax(0, 1fr));
  gap: 12px;
}

.profile-fact {
  display: grid;
  gap: 6px;
  min-height: 100%;
  padding: 16px 18px;
  border: 1px solid var(--line);
  border-radius: 12px;
  background: var(--white);
  box-shadow: 0 8px 20px rgba(20, 38, 28, 0.04);
}

.profile-fact-label {
  color: var(--copper);
  font-family: var(--font-sans);
  font-size: 0.72rem;
  font-weight: 800;
  letter-spacing: 0.06em;
  text-transform: uppercase;
}

.profile-fact-value {
  color: var(--ink);
  font-size: 0.98rem;
  font-weight: 600;
  line-height: 1.4;
}

.profile-recognize {
  margin-top: clamp(28px, 4vw, 40px);
  padding: clamp(20px, 3vw, 28px);
  border-radius: 14px;
  background: var(--white);
  border: 1px solid var(--line);
}

.profile-recognize h3 {
  margin: 0 0 12px;
  color: var(--forest);
}

.profile-list {
  max-width: 48rem;
  margin: 0;
  padding-left: 1.2rem;
  color: var(--muted);
  line-height: 1.7;
}

.profile-list li + li {
  margin-top: 10px;
}

.profile-list-rich li {
  padding-left: 4px;
}

.lookalike-grid {
  display: grid;
  grid-template-columns: repeat(3, minmax(0, 1fr));
  gap: 16px;
  margin-top: 8px;
}

.lookalike-card {
  display: grid;
  gap: 10px;
  align-content: start;
  padding: 20px;
  border: 1px solid var(--line);
  border-radius: 14px;
  background: var(--white);
  box-shadow: 0 10px 24px rgba(20, 38, 28, 0.05);
}

.lookalike-card h3 {
  display: flex;
  flex-wrap: wrap;
  align-items: center;
  gap: 8px;
  margin: 0;
  font-size: 1.2rem;
  line-height: 1.25;
}

.lookalike-card h3 a {
  text-decoration: none;
}

.lookalike-card h3 a:hover,
.lookalike-card h3 a:focus-visible {
  color: var(--copper);
}

.lookalike-card p {
  margin: 0;
  font-size: 0.98rem;
  line-height: 1.6;
}

.location-layout {
  display: grid;
  grid-template-columns: minmax(220px, 0.85fr) minmax(320px, 1.15fr);
  gap: clamp(28px, 5vw, 56px);
  align-items: center;
}

.location-copy h3 {
  margin: 20px 0 10px;
  color: var(--ink);
  font-size: 1rem;
}

.location-copy p {
  margin: 0 0 12px;
}

.profile-map-note {
  margin: 18px 0 0;
  color: var(--muted);
  font-size: 0.88rem;
}

.image-gallery {
  display: grid;
  grid-template-columns: repeat(2, minmax(0, 1fr));
  gap: clamp(18px, 3vw, 32px);
  max-width: 1120px;
  margin: 0 auto;
  padding: 8px clamp(18px, 4vw, 56px) 0;
}

.gallery-figure {
  margin: 0;
}

.gallery-figure picture {
  display: block;
  overflow: hidden;
  border-radius: 12px;
  background: var(--soft);
  border: 1px solid var(--line);
}

.gallery-image {
  display: block;
  width: 100%;
  height: auto;
  aspect-ratio: 4 / 3;
  object-fit: cover;
}

.gallery-figure figcaption {
  display: grid;
  gap: 4px;
  margin-top: 10px;
  color: var(--muted);
  font-size: 0.82rem;
  line-height: 1.45;
}

.gallery-caption {
  color: var(--ink);
  font-weight: 600;
}

.gallery-credit {
  font-size: 0.78rem;
}

.profile-section-sources .profile-resource-list {
  padding: 0;
}

@media (max-width: 900px) {
  .profile-fact-grid,
  .lookalike-grid {
    grid-template-columns: 1fr 1fr;
  }
}

@media (max-width: 700px) {
  .image-gallery {
    grid-template-columns: 1fr;
  }

  .profile-fact-grid,
  .lookalike-grid {
    grid-template-columns: 1fr;
  }
}

.range-map {
  margin: 0;
  padding: clamp(18px, 3vw, 28px);
  border: 1px solid var(--line);
  border-radius: 8px;
  background: var(--soft);
}

.range-map-graphic {
  display: block;
  width: 100%;
  height: auto;
}

.range-region {
  fill: rgba(255, 255, 255, 0.66);
}

.range-region.is-active {
  fill: var(--copper);
}

.range-boundaries {
  fill: none;
  stroke: var(--forest);
  stroke-linecap: round;
  stroke-linejoin: round;
  stroke-width: 2.5;
  vector-effect: non-scaling-stroke;
}

.range-region-label {
  fill: var(--muted);
  font-family: var(--font-sans);
  font-size: 17px;
  font-weight: 800;
  paint-order: stroke;
  stroke: rgba(255, 254, 251, 0.88);
  stroke-width: 4px;
  stroke-linejoin: round;
  text-anchor: middle;
}

.range-region-label.is-active {
  fill: var(--white);
  stroke: rgba(49, 63, 46, 0.52);
}

.profile-resource-list {
  display: grid;
  gap: 12px;
  max-width: 900px;
  margin: 0;
  padding: 0;
  list-style: none;
}

.profile-resource-list li {
  padding: 14px 0;
  border-bottom: 1px solid var(--line);
}

.profile-resource-list a {
  color: var(--moss-dark);
  font-weight: 700;
  text-underline-offset: 3px;
}

.articles-hero {
  padding: clamp(52px, 7vw, 96px) clamp(18px, 7vw, 110px);
  background:
    radial-gradient(circle at 88% 14%, rgba(181, 106, 58, 0.32), transparent 32%),
    linear-gradient(110deg, rgba(20, 38, 28, 0.98), rgba(26, 51, 40, 0.88)),
    var(--forest);
  color: var(--white);
}

.articles-hero > div {
  max-width: 1050px;
}

.articles-hero h1 {
  max-width: 900px;
  margin: 0;
  font-size: clamp(3rem, 8vw, 4.5rem);
  letter-spacing: -0.05em;
  line-height: 0.94;
}

.articles-hero p {
  max-width: 760px;
  margin: 30px 0 0;
  color: rgba(255, 255, 255, 0.76);
  font-size: clamp(1.1rem, 2vw, 1.35rem);
  line-height: 1.65;
}

.article-series-section,
.more-guides {
  padding: clamp(52px, 8vw, 104px) clamp(18px, 7vw, 110px);
}

.article-series-section + .article-series-section {
  border-top: 1px solid var(--line);
  background: rgba(216, 226, 212, 0.42);
}

.series-heading {
  display: grid;
  grid-template-columns: minmax(0, 1fr) minmax(240px, 0.65fr);
  gap: 32px;
  align-items: end;
  max-width: 1180px;
  margin-bottom: clamp(32px, 5vw, 58px);
}

.series-heading h2,
.more-guides h2 {
  margin: 0;
  font-size: clamp(2.2rem, 5vw, 4.75rem);
  letter-spacing: -0.04em;
  line-height: 1;
}

.series-heading p {
  margin: 0;
  color: var(--muted);
  font-size: 1.05rem;
  line-height: 1.65;
}

.article-card-list {
  max-width: 1180px;
  margin: 0;
  padding: 0;
  border-top: 1px solid var(--line);
  list-style: none;
}

.article-card-list li {
  border-bottom: 1px solid var(--line);
}

.article-card-list a {
  display: grid;
  grid-template-columns: 72px minmax(0, 1fr);
  gap: clamp(18px, 4vw, 52px);
  padding: clamp(26px, 4vw, 48px) 0;
  text-decoration: none;
}

.article-card-list a:hover h3,
.article-card-list a:focus-visible h3 {
  color: var(--copper);
}

.article-card-number {
  padding-top: 5px;
  color: var(--copper);
  font-size: 0.9rem;
  font-weight: 800;
}

.article-card-list h3 {
  margin: 0;
  font-size: clamp(1.65rem, 3.5vw, 3.25rem);
  letter-spacing: -0.035em;
  line-height: 1.05;
  transition: color 160ms ease;
}

.article-card-list p {
  max-width: 790px;
  margin: 14px 0 0;
  color: var(--muted);
  line-height: 1.65;
}

.article-card-meta {
  display: block;
  margin-top: 16px;
  color: var(--muted);
  font-size: 0.82rem;
  font-weight: 700;
}

.more-guides {
  border-top: 1px solid var(--line);
  background: var(--sage);
}

.more-guides > a {
  display: grid;
  gap: 8px;
  max-width: 1180px;
  margin-top: 32px;
  padding: clamp(26px, 4vw, 42px) 0;
  border-block: 1px solid rgba(20, 38, 28, 0.18);
  text-decoration: none;
}

.more-guides strong {
  font-size: clamp(1.35rem, 3vw, 2.5rem);
}

.more-guides span {
  color: var(--muted);
  line-height: 1.55;
}

.behavior-hero {
  padding: clamp(52px, 7vw, 96px) clamp(18px, 7vw, 110px);
  background:
    radial-gradient(circle at 88% 12%, rgba(181, 106, 58, 0.36), transparent 32%),
    linear-gradient(145deg, #14261C, var(--forest));
  color: var(--white);
}

.behavior-hero-inner {
  max-width: 1050px;
}

.series-back-link {
  display: inline-block;
  color: #e8c4a0;
  font-family: var(--font-sans);
  font-size: 0.88rem;
  font-weight: 800;
  text-decoration-thickness: 1px;
  text-underline-offset: 4px;
}

.article-meta {
  margin: 28px 0 0;
  color: rgba(255, 255, 255, 0.62);
  font-size: 0.84rem;
  font-weight: 700;
}

.behavior-hero h1 {
  max-width: 920px;
  margin: 18px 0 0;
  font-size: clamp(3rem, 8vw, 4.5rem);
  letter-spacing: -0.05em;
  line-height: 0.94;
}

.article-subtitle {
  margin: 18px 0 0;
  color: #e8c4a0;
  font-family: var(--font-serif);
  font-size: clamp(1.15rem, 2.4vw, 1.65rem);
  font-weight: 600;
}

.behavior-hero .article-deck {
  max-width: 760px;
}

.article-layout {
  display: grid;
  grid-template-columns: minmax(0, 760px) minmax(260px, 340px);
  gap: clamp(48px, 8vw, 108px);
  align-items: start;
  max-width: 1280px;
  margin: 0 auto;
  padding: clamp(52px, 8vw, 104px) clamp(18px, 5vw, 70px);
}

.behavior-article {
  min-width: 0;
}

.article-lede {
  padding-bottom: clamp(28px, 4vw, 48px);
  border-bottom: 1px solid var(--line);
}

.article-lede p,
.behavior-article section > p {
  margin: 0 0 24px;
  color: var(--muted);
  font-size: clamp(1.05rem, 1.6vw, 1.18rem);
  line-height: 1.82;
}

.behavior-article > section {
  margin-top: clamp(48px, 7vw, 78px);
}

.behavior-article h2 {
  margin: 0 0 24px;
  font-size: clamp(1.9rem, 4vw, 3rem);
  letter-spacing: -0.035em;
  line-height: 1.08;
}

.field-note {
  margin-top: clamp(48px, 7vw, 78px);
  padding: clamp(26px, 5vw, 44px);
  border-left: 5px solid var(--copper);
  background: var(--sage);
}

.field-note h2 {
  margin: 0 0 10px;
  color: var(--copper);
  font-size: 0.86rem;
  letter-spacing: 0;
  text-transform: uppercase;
}

.field-note p {
  margin: 0;
  font-size: clamp(1.05rem, 2vw, 1.22rem);
  line-height: 1.7;
}

.article-sources {
  padding-top: clamp(42px, 6vw, 66px);
  border-top: 1px solid var(--line);
}

.article-sources h2 {
  font-size: clamp(1.45rem, 3vw, 2rem);
}

.article-sources ul {
  display: grid;
  gap: 12px;
  margin: 0;
  padding-left: 20px;
  color: var(--muted);
  line-height: 1.55;
}

.article-sources a {
  font-weight: 700;
  text-underline-offset: 3px;
}

.series-sidebar {
  position: sticky;
  top: 118px;
  padding-top: 4px;
}

.series-sidebar > h2 {
  margin: 0 0 16px;
  font-size: 1rem;
}

.series-list {
  margin: 0;
  padding: 0;
  border-top: 1px solid var(--line);
  list-style: none;
}

.series-list li {
  border-bottom: 1px solid var(--line);
}

.series-list a {
  display: grid;
  grid-template-columns: 28px minmax(0, 1fr);
  gap: 10px;
  padding: 16px 0;
  color: var(--muted);
  text-decoration: none;
}

.series-list span {
  color: var(--copper);
  font-size: 0.75rem;
  font-weight: 800;
}

.series-list strong {
  font-size: 0.9rem;
  line-height: 1.35;
}

.series-list .is-current a {
  color: var(--ink);
}

.article-pagination {
  display: grid;
  grid-template-columns: repeat(2, minmax(0, 1fr));
  gap: 1px;
  margin-top: clamp(50px, 8vw, 90px);
  padding: 1px;
  background: var(--line);
}

.article-pagination a {
  display: grid;
  align-content: start;
  gap: 8px;
  min-height: 128px;
  padding: 24px;
  background: var(--white);
  text-decoration: none;
}

.article-pagination .next {
  text-align: right;
}

.article-pagination span {
  color: var(--muted);
  font-size: 0.78rem;
  font-weight: 700;
}

.article-pagination strong {
  line-height: 1.4;
}

@media (max-width: 1120px) {
  .species-grid {
    grid-template-columns: repeat(3, minmax(0, 1fr));
  }

  .profile-hero {
    grid-template-columns: 1fr;
  }

  .location-layout {
    grid-template-columns: 1fr;
  }

  .article-layout {
    grid-template-columns: minmax(0, 760px);
    justify-content: center;
  }

  .series-sidebar {
    position: static;
    order: -1;
  }

}

@media (max-width: 780px) {
  .site-header {
    min-height: 76px;
    padding-block: 12px;
  }

  .brand {
    gap: 10px;
  }

  .brand-text {
    font-size: 1.28rem;
  }

  .brand-mark {
    width: 40px;
    height: 40px;
  }

  .site-nav {
    gap: 16px;
  }

  .site-nav a {
    font-size: 0.88rem;
  }

  .home-articles {
    align-items: start;
    flex-direction: column;
  }

  .home-articles h1,
  .articles-hero h1,
  .behavior-hero h1 {
    font-size: clamp(2.35rem, 12vw, 3.4rem);
  }

  .series-heading {
    grid-template-columns: 1fr;
  }

  .series-sidebar {
    order: initial;
  }

  .article-card-list a {
    grid-template-columns: 42px minmax(0, 1fr);
  }

  .article-pagination {
    grid-template-columns: 1fr;
  }

  .article-pagination .next {
    text-align: left;
  }

  .species-grid {
    grid-template-columns: 1fr;
  }

  .species-search-row {
    grid-template-columns: 1fr;
    gap: 8px;
  }

  .species-search output {
    text-align: left;
  }

  .species-compare-grid {
    grid-template-columns: 1fr;
  }

}

@media (max-width: 520px) {
  .site-header {
    align-items: flex-start;
    flex-direction: column;
    gap: 4px;
  }

  .site-nav a {
    padding-block: 7px;
  }
}